Chapter 5 · 23 min

Perceiving, Learning, and Remembering

Chapter 5 of How the Brain Works: Perceiving, Learning, and Remembering.

How the Brain Works | Ch 5 Part 1/2: Perceiving, Learning, and Remembering ↗How the Brain Works | Ch 5 Part 2/2: Perceiving, Learning, and Remembering ↗

Visual credits

7 sources

Sources and credits for imagery used in this film, including material incorporated into explanatory animations.

  • Subdural grid on cortical surface, original PLOS Figure2

    Edwin van Dellen, Linda Douw, Johannes C. Baayen, Jan J. Heimans, Sophie C. Ponten, W. Peter Vandertop, Demetrios N. Velis, Cornelis J. Stam, Jaap C. Reijneveld

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Cajal hippocampal circuit drawing, modified and labeled

    Santiago Ramon y Cajal; derivative labels and inset by Looie496

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Cultured cortical neuron with dendritic spines

    Sergb95

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Live confocal time-lapse of stable and retracting hippocampal dendritic spines

    J. Reilly, H. Hanson, M. Fernandez-Monreal, S. Wearne, P. Hof, and G. Phillips

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Woollett and Maguire 2011 Figure 2: posterior hippocampal gray-matter change

    Katherine Woollett and Eleanor A. Maguire

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Brenda Milner, 2014 portrait

    Per Henning/NTNU

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

  • Roy2022 brain-wide activity-dependent maps, Figure1

    Dheeraj S. Roy and coauthors

    Incorporated into an explanatory animation; presentation may include cropping, labels and overlays.

Research references

Selected sources from the episode’s research and scientific figures.

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  2. DOI 10.1113/jphysiol.1973.sp010273 ↗
  3. author PDF ↗
  4. Corkin, 2002 ↗
  5. https://www.nature.com/articles/ncomms4122 ↗
  6. Nature Communications 13, 1799 ↗
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  8. Talarico and Rubin, Psychological Science 2003, DOI 10.1111/1467-9280.02453 ↗
  9. Collingridge 1983 ↗
  10. Dudek/Bear 1992 ↗
  11. Saucier/Cain 1995 ↗
  12. Nature DOI 10.1038/nature08389 ↗
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  14. PubMed abstract ↗
  15. Nature Neuroscience DOI 10.1038/s41593-026-02230-2 ↗
  16. primary abstract ↗
  17. complete Edinburgh author manuscript ↗
  18. primary paper ↗
  19. Figure 1 ↗
  20. PLOS ONE Figure 2 ↗
  21. “Brenda Milner – Neuropsychology,” 4:03 ↗
  22. Science Museum Group wooden mirror-drawing apparatus, object 1999-1027 ↗
  23. MNCN collection ↗
  24. HHMI BioInteractive “Spiny Cells” page ↗
  25. PMC reference copy ↗
  26. Bliss and Gardner-Medwin, 1973, companion primary paper ↗
  27. Collingridge, Kehl and McLennan, 1983 ↗
  28. historical account ↗
  29. Morris, Anderson, Lynch and Baudry, 1986 ↗
  30. Saucier and Cain, 1995 ↗
  31. Dudek and Bear, 1992 ↗
  32. mechanism reference ↗
  33. Nicoll 2017 review ↗
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  36. publisher and extended-data captions ↗
  37. PMC identity ↗
  38. Sian Lewis commentary, “Engineering memories” ↗
  39. “The Role of Synapses in Memory – On Our Mind,” 16:24 ↗
  40. PMC copy ↗
  41. Investigator institution's account ↗
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  43. Investigator institutional summary ↗
  44. Primary article ↗
  45. UCL PDF ↗
  46. https://www.ebi.ac.uk/europepmc/webservices/rest/PMC3268356/fullTextXML ↗
  47. PMC497229 ↗
  48. university-hosted scan ↗
  49. UCSD ↗
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  51. MIT 2013 interview ↗
  52. MIT's 2016 statement ↗
  53. Seth Mnookin's contemporaneous account ↗
  54. Popular Science ↗
  55. Milner lecture excerpt, 14:08 ↗
  56. https://doi.org/10.1016/0028-3932(68)90024-9 ↗
  57. Nature DOI 10.1038/nature11028 ↗
  58. investigators' methods account ↗
  59. Science DOI 10.1126/science.aam6808 ↗
  60. Lab PDF ↗
  61. MIT image ↗
  62. Science DOI 10.1126/science.ado8316 ↗
  63. author manuscript PMC12233322 ↗
  64. 2024 preprint record ↗
  65. NIMH reconstruction images ↗
  66. PubMed ↗
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  88. primary article ↗
  89. primary PDF ↗
  90. primary abstract ↗
  91. primary paper ↗
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  94. NCBI Neuroscience chapter NBK10941 ↗
  95. Castellucci et al., Science 1970 ↗
  96. Castellucci and Kandel, PNAS 1974 ↗
  97. Kandel's Nobel lecture ↗
  98. Nobel organization record ↗
  99. Kandel's first-person scientific biography ↗
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  105. DOI 10.1136/jnnp.20.1.11 ↗
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  107. Nature DOI 10.1038/nature13294 ↗
  108. DOI 10.1016/j.cub.2011.11.018 ↗
  109. official reproduction conditions ↗
  110. specific Simurg item ↗
  111. 2014 UCSD report ↗
  112. Commons ↗
  113. https://news.mit.edu/2013/suzanne-corkin-permanent-present-tense-0514): ↗
  114. https://today.ucsd.edu/story/h.m.s_brain_yields_new_evidence): ↗
  115. https://www.youtube.com/watch?v=VYNh3-21HmA): ↗
  116. https://www.youtube.com/watch?v=GhfNtyL_Cj0): ↗
  117. https://www.youtube.com/watch?v=NehVPiKJNqs): ↗

How this film was made

Created and editorially directed by Anders Lunde using a custom production system developed over six months. AI assists research, drafting, programming, and visual production; I review and edit the material and make the final editorial decisions. Narration uses text-to-speech. Visuals combine custom-built scientific animations, AI-generated reconstructions, and credited scientific imagery.

Generated reconstructions illustrate scientific interpretations. They are distinct from photographs, specimens, and measured data.

More about the creator and production →